I've re enabled full UAC on my main PC to check again.
UAC does not prevent saving from any web browser, neither does it prevent right-click Save Target As... or Save Picture As...
There may be some additional 3rd party security software on Chris' PC which is closely tied to UAC and offers additional safety features not normally shipped with a standard Vista installation.
There's a handy UAC utility available here which enables 'Quiet mode' for those who want the additional protection but prefer to suppress the elevation prompts for administrators .
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